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Anatomical Evidence for an Indirect Pathway for Word Repetition

Neurology 2020 

Watch my talk on Youtube: 



Book Chapters &

other media

Forkel SJ et al. Clinical applications of diffusion tractography. In the Handbook of Diffusion Imaging. (Dell’Acqua & Leemanns, Eds), forthcoming


Forkel SJ. White matter variability, cognition and disorders. IIn: Della Sala, S. (Ed.), Encyclopedia of Behavioral Neuroscience, vol. 2. Elsevier, pp. 233–241. https://dx.doi.org/10.1016/B978-0-12-819641-0.00086-4.


Forkel SJ. Lesion-symptom mapping: from single cases to the human disconnectome. In Encyclopaedia of Behavioural Neuroscience 2e, (Della Sala et al., Eds). Elsevier, 10.13140/RG.2.2.26163.66088


Catani M & Forkel SJ. Diffusion Imaging Methods in Language Sciences. In Oxford Handbook of Neurolinguistics (de Zubicaray & Schiller, Eds). OUP, 2019


Forkel SJ & Catani M. Structural Neuroimaging. In Research Methods in Psycholinguistics (de Groot & Hoogart, Eds). Wiley & Sons, 2018


Rickards T, Baldo J, Yochim BP. Changes to the Brain: Methods of Investigation, Aging, and Neuroplasticity (image contribution) in Yochim & Woodhead (Eds). Psychology of Aging: A Biopsychosocial Perspective. Springer, 2017

Forkel SJ & Howells H. Professor Blackburn on Unconscious biases. Published online on the Lindau Nobel Laureate website, 2015

Design Patent.  Design accessible via EUIPO Registered Community Design no.003823012-0001


Catani M, Forkel SJ, Thiebaut de Schotten M. Asymmetry of white matter pathways in the brain. In The Two Halves Of The Brain: Information Processing In The Cerebral Hemispheres (Eds, Hugdahl, K & Davidson, RJ). MIT Press, 2010

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Encyclopedia of Behavioral Neuroscience, second edition (BEV2)


My research follows several lines of investigation: 

  • Mapping neurovariability to predict clinical impairments and recovery for precision medicine

  • Anatomical studies of the human brain to map and validate the white matter networks

  • Comparative Anatomy studies to map the evolution of the brain as the foundation of higher cognitive functions

F1000 recommendations:


Boraud Th & Forkel SJ. Paul Broca: from fame to shame? Brain, in press. DOI: ​10.13140/RG.2.2.34092.59527

Mekki YN, Guillemot V, Lemaître H, Carrion-Castillo A, Forkel SJ, Frouin V, Philippe C. The genetic architecture of language functional connectivity, under revision, preprint : https://doi.org/10.1101/2021.10.18.464351 

Akeret, K, Forkel SJ,  Buzzi RM, Vasella F, Amrein I, Slomianka L, Serra C, Yasargil G, Krayenbühl N. The complex history of the forniceal commissure in a new light: a multimodal anatomy study, in prep

Talozzi L, Forkel SJ, Pacella V, Nozais V, Nachev P, Corbetta M, Thiebaut De Schotten M. Large scale disconnectome prediction of individual symptom severity in chronic stroke, 10.13140/RG.2.2.17963.36641

Watch the presentation here: https://youtu.be/sivOs4Y-YQI

Dulyan L, Talozzi L,  Pacella V,  Corbetta M,  Forkel SJ*, Thiebaut de Schotten* MLongitudinal prediction of motor dysfunction after stroke: a disconnectome study, in prep *joint last


White matter variability, cognition, and disorders: a systematic review.

Forkel SJ, Friedrich P, Thiebaut de Schotten M, Howells H.

Accepted in BSAF, 10.1007/s00429-021-02382-w 

Functionnectome: a framework to analyse the contribution of brain circuits to fMRI.

Nozais V, Forkel SJ, Petit L, Thiebaut de Schotten M.

 Communications Biology, | https://doi.org/10.1038/s42003-021-02530-2 

Towards metabolic disconnection – symptom mapping

Forkel SJ & Thiebaut de Schotten

Brain, doi: 10.1093/brain/awaa060, 2020

The effect of fronto-parietal resection on hand selection for grasping and manipulation

Howells HR, Puglisi G, Leonetti A, Vigano L, Fornia L, SimoneL, Forkel SJ, Rossi M, Cerri G, Bello L.

Cortex, 2020


Anatomical evidence of an indirect pathway for word repetition.

Forkel SJ, Rogalski E, Drossinos Sancho N, D'Anna L, Luque Laguna P, Sridhar J, Dell'Acqua F, Weintraub S, Thompson C, Catani M & Mesulam M-M

Neurology, 2020, F1000 Prime: https://f1000.com/prime/737290765

One size fits all does not apply to brain lateralisation.

Thiebaut de Schotten M, Friedrich P, Forkel SJ.

Physics of Life Reviews, doi: 10.1016/j.plrev.2019.07.007, 2019


Lesion mapping in acute stroke aphasia and its implications for recovery.

Forkel SJ & Catani M.

Neuropsychologia 115:88-100, 2018

Anatomical predictors of aphasia recovery: a tractography study of bilateral perisylvian language networks.

Forkel SJ, Thiebaut de Schotten M, Dell'Acqua F, Kalra L, Murphy, Williams S & Catani.

Brain 137(Pt 7):2027-39, 2014, F1000 Prime: https://facultyopinions.com/prime/718472585

Beyond cortical localization in clinico-anatomical correlation.

Catani M, Dell'Acqua F, Bizzi A, Forkel SJ, Williams SC, Simmons A, Murphy DG & Thiebaut de Schotten M.

Cortex 48(10):1262-87, 2012


A lateralized brain network for visuospatial attention.

Thiebaut de Schotten M, Dell'Acqua F, Forkel SJ, Simmons A, Vergani F, Murphy DG & Catani M.

Nature Neuroscience 14(10):1245-6, 2011


Altered integrity of perisylvian language pathways in schizophrenia: relationship to auditory hallucinations.

Catani M, Craig MC, Forkel SJ, Kanaan R, Picchioni M, Toulopoulou T, Shergill S, Williams S, Murphy DG & McGuire P.

Biological Psychiatry 70(12):1143-50, 2011



Anchoring the human olfactory system to a functional gradient.

Waymel A, Friedrich P, Forkel SJ*, Thiebaut de Schotten M*. *joint last.

NeuroImage, doi: 10.1016/j.neuroimage.2020.116863, 2020

Mapping the principal gradient onto the corpus callosum

Friedrich P, Forkel SJ, Thiebaut de Schotten M.

NeuroImage, doi: https://doi.org/10.1101/2020.03.05.978171, 2020

An ancestral anatomical and spatial bias for visually guided behavior

Friedrich P, Thiebaut de Schotten M, Forkel SJ, Stacho M, Howells HR

PNAS, 117(5):2251-2252. doi: 10.1073/pnas.1918402117, 2020

The white matter of the human cerebrum: part I The occipital lobe by Heinrich Sachs.

Forkel SJ, Mahmood S, Vergani F & Catani M.

Cortex 62:182-202, 2015

Intralobar fibres of the occipital lobe: a post mortem dissection study.

Vergani F, Mahmood S, Morris CM, Mitchell P & Forkel SJ.

Cortex 56:145-56, 2014

The anatomy of fronto-occipital connections from early blunt dissections to contemporary tractography.

Forkel SJ, Thiebaut de Schotten M, Kawadler JM, Dell'Acqua F, Danek A & Catani M.

Cortex 56:73-84, 2014


Saimiri model in Clinical Neuroscience Research

Royo J, Forkel SJ, Pouget P, Thiebaut de Schotten M. 

Neuroscience & Biobehavioral Reviews, accepted. DOI: 10.13140/RG.2.2.34359.55200/1.

Imaging the primate brain evolution: the next frontier?

Friedrich P, Forkel SJ, Amiez C, Balsters JH, Coulon O, Fan L, Goulas A, Hadj-Bouziane F, Hecht EE, Heuer K, Jiang TLatzman RD, Liu X , Kee Loh K, Patil KR, Lopez-Persem A, Procyk E, Sallet J, Toro R, Vickery S, Weis S, Wilson C, Xu T, Zerbi V, Eickoff SB *, Margulies D*, Mars RB*, Thiebaut de Schotten M* joint last

NeuroImage, https://doi.org/10.1016/j.neuroimage.2020.117685, 2021


Accelerating the evolution of nonhuman primate neuroimaging 

Milham M, Petkov C, Margulies M, Schroeder C, Basso MA, Belin P, Fox A, Kastner S, Mars RB, Messinger A, Poirier C, Vanduffel W, Van Essen DC, Albin S, Alvand A, Becker Y, Ben Hamed S, Benn A, Bodin C, Boretius S, Cagna B, Coulon O, ElGohary SH, Evrard H, Forkel SJ,

Friedrich P, Froudist-Walsh S, Gao Y, Gozzi A, Grigis A, Hartig R, Hayashi T, Solon Heinsfeld A, Heuer K, Howells Z, Jan Ardesch D, Kagan I, Kelly C, Kennedy H, Klink C, Konen C, Chai Kwok S, Leech R, Liu X, Madan C, Madushanka W, Majka P, Mallon AM, Marche K, Meguerditchian A, Menon R, Merchant H, Messinger A, Mitchell A, Nenning KH, Nikolaidis A, Ortiz-Rios M, Pagani M, Pareek V, Petkov C, Pool RR, Prescott M, Procyk E, Rajimehr R, Rautu IS, Raz A, Roe AW, Roumazeilles L, Sakai T, Sallet J, Saldivar PJ, Sato C, Sawiak S, Schiffer M, Schroeder CE, Schwiedrzik CM, Seidlitz J, Sein J, Shen Z, Silva A, Simone L, Sirmpilatze N, Sliwa J, Smallwood J, Tasserie J, Toro R, Thiebaut de Schotten M, Trapeau R, Vezoli J, Garza Villarreal E, Wang Z, Wells S, Williams B, Xu T, Xu AG, Yacoub E, and Zhan M

Neuron, 105(4):600-603. doi: 10.1016/j.neuron.2019.12.023, 2020


Differences in frontal network anatomy across primate species.

Barrett R, Dawson M, Dyrby T, Krug K, Ptito M, D'Arceuil H, Croxson PL, Johnson P, Howells H, Forkel SJ, Dell’Acqua F & Catani M

Journal of Neuroscience, 2020

Structural Variability Across the Primate Brain: A Cross-Species Comparison. Croxson PL, Forkel SJ, Cerliani L & Thiebaut de Schotten M

Cereb Cortex 28(11):3829-3841, 2018

Short parietal lobe connections of the human and monkey brain.

Catani M, Robertsson N, Beyh A, Huynh V, de Santiago Requejo F, Howells H, Barrett RL, Aiello M, Cavaliere C, Dyrby T, Krug K, Ptito M, D'Arceuil H & Forkel SJ*, Dell'Acqua F*. *joint last.

Cortex 97:339-357, 2017